Regulation of neuronal axon specification by glia-neuron gap junctions in <i>C. elegans</i>.

Meng, Lingfeng; Zhang, Albert; Jin, Yishi; Yan, Dong · Elife · 2016

basic_science · Level V

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Abstract

Axon specification is a critical step in neuronal development, and the function of glial cells in this process is not fully understood. Here, we show that <i>C. elegans</i> GLR glial cells regulate axon specification of their nearby GABAergic RME neurons through GLR-RME gap junctions. Disruption of GLR-RME gap junctions causes misaccumulation of axonal markers in non-axonal neurites of RME neurons and converts microtubules in those neurites to form an axon-like assembly. We further uncover that GLR-RME gap junctions regulate RME axon specification through activation of the CDK-5 pathway in a calcium-dependent manner, involving a calpain <i>clp-4</i>. Therefore, our study reveals the function of glia-neuron gap junctions in neuronal axon specification and shows that calcium originated from glial cells can regulate neuronal intracellular pathways through gap junctions.

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