<i>bicoid</i> mRNA localises to the <i>Drosophila</i> oocyte anterior by random Dynein-mediated transport and anchoring.

Trovisco, Vítor; Belaya, Katsiaryna; Nashchekin, Dmitry; Irion, Uwe; Sirinakis, George; Butler, Richard; Lee, Jack J; Gavis, Elizabeth R et al. · Elife · 2016

basic_science · Level V

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Abstract

<i>bicoid</i> mRNA localises to the <i>Drosophila</i> oocyte anterior from stage 9 of oogenesis onwards to provide a local source for Bicoid protein for embryonic patterning. Live imaging at stage 9 reveals that <i>bicoid</i> mRNA particles undergo rapid Dynein-dependent movements near the oocyte anterior, but with no directional bias. Furthermore, <i>bicoid</i> mRNA localises normally in <i>shot<sup>2A2</sup></i>, which abolishes the polarised microtubule organisation. FRAP and photo-conversion experiments demonstrate that the RNA is stably anchored at the anterior, independently of microtubules. Thus, <i>bicoid</i> mRNA is localised by random active transport and anterior anchoring. Super-resolution imaging reveals that <i>bicoid</i> mRNA forms 110-120 nm particles with variable RNA content, but constant size. These particles appear to be well-defined structures that package the RNA for transport and anchoring.

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