Structural dynamics of RbmA governs plasticity of <i>Vibrio cholerae</i> biofilms.

Fong, Jiunn Cn; Rogers, Andrew; Michael, Alicia K; Parsley, Nicole C; Cornell, William-Cole; Lin, Yu-Cheng; Singh, Praveen K; Hartmann, Raimo et al. · Elife · 2017

basic_science · Level V

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Abstract

Biofilm formation is critical for the infection cycle of <i>Vibrio cholerae. Vibrio</i> exopolysaccharides (VPS) and the matrix proteins RbmA, Bap1 and RbmC are required for the development of biofilm architecture. We demonstrate that RbmA binds VPS directly and uses a binary structural switch within its first fibronectin type III (FnIII-1) domain to control RbmA structural dynamics and the formation of VPS-dependent higher-order structures. The structural switch in FnIII-1 regulates interactions in trans with the FnIII-2 domain, leading to open (monomeric) or closed (dimeric) interfaces. The ability of RbmA to switch between open and closed states is important for <i>V. cholerae</i> biofilm formation, as RbmA variants with switches that are locked in either of the two states lead to biofilms with altered architecture and structural integrity.

Medical subject headings