Structure of a Wbl protein and implications for NO sensing by M. tuberculosis.

Kudhair, Bassam K; Hounslow, Andrea M; Rolfe, Matthew D; Crack, Jason C; Hunt, Debbie M; Buxton, Roger S; Smith, Laura J; Le Brun, Nick E et al. · Nat Commun · 2017

basic_science · Level V

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Abstract

Mycobacterium tuberculosis causes pulmonary tuberculosis (TB) and claims ~1.8 million human lives per annum. Host nitric oxide (NO) is important in controlling TB infection. M. tuberculosis WhiB1 is a NO-responsive Wbl protein (actinobacterial iron-sulfur proteins first identified in the 1970s). Until now, the structure of a Wbl protein has not been available. Here a NMR structural model of WhiB1 reveals that Wbl proteins are four-helix bundles with a core of three α-helices held together by a [4Fe-4S] cluster. The iron-sulfur cluster is required for formation of a complex with the major sigma factor (σ<sup>A</sup>) and reaction with NO disassembles this complex. The WhiB1 structure suggests that loss of the iron-sulfur cluster (by nitrosylation) permits positively charged residues in the C-terminal helix to engage in DNA binding, triggering a major reprogramming of gene expression that includes components of the virulence-critical ESX-1 secretion system.

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