Thioredoxin shapes the <i>C. elegans</i> sensory response to <i>Pseudomonas</i> produced nitric oxide.

Hao, Yingsong; Yang, Wenxing; Ren, Jing; Hall, Qi; Zhang, Yun; Kaplan, Joshua M · Elife · 2018

basic_science · Level V

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Abstract

Nitric oxide (NO) is released into the air by NO-producing organisms; however, it is unclear if animals utilize NO as a sensory cue. We show that <i>C. elegans</i> avoids <i>Pseudomonas aeruginosa</i> (PA14) in part by detecting PA14-produced NO. PA14 mutants deficient for NO production fail to elicit avoidance and NO donors repel worms. PA14 and NO avoidance are mediated by a chemosensory neuron (ASJ) and these responses require receptor guanylate cyclases and cyclic nucleotide gated ion channels. ASJ exhibits calcium increases at both the onset and removal of NO. These NO-evoked ON and OFF calcium transients are affected by a redox sensing protein, TRX-1/thioredoxin. TRX-1's trans-nitrosylation activity inhibits the ON transient whereas TRX-1's de-nitrosylation activity promotes the OFF transient. Thus, <i>C. elegans</i> exploits bacterially produced NO as a cue to mediate avoidance and TRX-1 endows ASJ with a bi-phasic response to NO exposure.

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