The adhesion function of the sodium channel beta subunit (β1) contributes to cardiac action potential propagation.

Veeraraghavan, Rengasayee; Hoeker, Gregory S; Alvarez-Laviada, Anita; Hoagland, Daniel; Wan, Xiaoping; King, D Ryan; Sanchez-Alonso, Jose; Chen, Chunling et al. · Elife · 2018

basic_science · Level V

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Abstract

Computational modeling indicates that cardiac conduction may involve ephaptic coupling - intercellular communication involving electrochemical signaling across narrow extracellular clefts between cardiomyocytes. We hypothesized that β1(SCN1B) -mediated adhesion scaffolds <i>trans</i>-activating Na<sub>V</sub>1.5 (SCN5A) channels within narrow (<30 nm) perinexal clefts adjacent to gap junctions (GJs), facilitating ephaptic coupling. Super-resolution imaging indicated preferential β1 localization at the perinexus, where it co-locates with Na<sub>V</sub>1.5. Smart patch clamp (SPC) indicated greater sodium current density (I<sub>Na</sub>) at perinexi, relative to non-junctional sites. A novel, rationally designed peptide, βadp1, potently and selectively inhibited β1-mediated adhesion, in electric cell-substrate impedance sensing studies. βadp1 significantly widened perinexi in guinea pig ventricles, and selectively reduced perinexal I<sub>Na</sub>, but not whole cell I<sub>Na</sub>, in myocyte monolayers. In optical mapping studies, βadp1 precipitated arrhythmogenic conduction slowing. In summary, β1-mediated adhesion at the perinexus facilitates action potential propagation between cardiomyocytes, and may represent a novel target for anti-arrhythmic therapies.

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