Rbpj expression in regulatory T cells is critical for restraining T<sub>H</sub>2 responses.

Delacher, Michael; Schmidl, Christian; Herzig, Yonatan; Breloer, Minka; Hartmann, Wiebke; Brunk, Fabian; Kägebein, Danny; Träger, Ulrike et al. · Nat Commun · 2019

basic_science · Level V

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Abstract

The transcriptional regulator Rbpj is involved in T-helper (T<sub>H</sub>) subset polarization, but its function in T<sub>reg</sub> cells remains unclear. Here we show that T<sub>reg</sub>-specific Rbpj deletion leads to splenomegaly and lymphadenopathy despite increased numbers of T<sub>reg</sub> cells with a polyclonal TCR repertoire. A specific defect of Rbpj-deficient T<sub>reg</sub> cells in controlling T<sub>H</sub>2 polarization and B cell responses is observed, leading to the spontaneous formation of germinal centers and a T<sub>H</sub>2-associated immunoglobulin class switch. The observed phenotype is environment-dependent and can be induced by infection with parasitic nematodes. Rbpj-deficient T<sub>reg</sub> cells adopt open chromatin landscapes and gene expression profiles reminiscent of tissue-derived T<sub>H</sub>2-polarized T<sub>reg</sub> cells, with a prevailing signature of the transcription factor Gata-3. Taken together, our study suggests that T<sub>reg</sub> cells require Rbpj to specifically restrain T<sub>H</sub>2 responses, including their own excessive T<sub>H</sub>2-like differentiation potential.

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