The Fml1-MHF complex suppresses inter-fork strand annealing in fission yeast.

Wong, Io Nam; Neo, Jacqueline Ps; Oehler, Judith; Schafhauser, Sophie; Osman, Fekret; Carr, Stephen B; Whitby, Matthew C · Elife · 2019

basic_science · Level V

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Abstract

Previously we reported that a process called inter-fork strand annealing (IFSA) causes genomic deletions during the termination of DNA replication when an active replication fork converges on a collapsed fork (Morrow et al., 2017). We also identified the FANCM-related DNA helicase Fml1 as a potential suppressor of IFSA. Here, we confirm that Fml1 does indeed suppress IFSA, and show that this function depends on its catalytic activity and ability to interact with Mhf1-Mhf2 via its C-terminal domain. Finally, a plausible mechanism of IFSA suppression is demonstrated by the finding that Fml1 can catalyse regressed fork restoration in vitro.

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