Long non-coding RNA <i>GRASLND</i> enhances chondrogenesis via suppression of the interferon type II signaling pathway.

Huynh, Nguyen Pt; Gloss, Catherine C; Lorentz, Jeremiah; Tang, Ruhang; Brunger, Jonathan M; McAlinden, Audrey; Zhang, Bo; Guilak, Farshid · Elife · 2020

basic_science · Level V

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Abstract

The roles of long noncoding RNAs (lncRNAs) in musculoskeletal development, disease, and regeneration remain poorly understood. Here, we identified the novel lncRNA <i>GRASLND</i> (originally named <i>RNF144A-AS1</i>) as a regulator of mesenchymal stem cell (MSC) chondrogenesis. <i>GRASLND</i>, a primate-specific lncRNA, is upregulated during MSC chondrogenesis and appears to act directly downstream of SOX9, but not TGF-β3. We showed that the silencing of <i>GRASLND</i> resulted in lower accumulation of cartilage-like extracellular matrix in a pellet assay, while <i>GRASLND</i> overexpression - either via transgene ectopic expression or by endogenous activation via CRISPR-dCas9-VP64 - significantly enhanced cartilage matrix production. <i>GRASLND</i> acts to inhibit IFN-γ by binding to EIF2AK2, and we further demonstrated that <i>GRASLND</i> exhibits a protective effect in engineered cartilage against interferon type II. Our results indicate an important role of <i>GRASLND</i> in regulating stem cell chondrogenesis, as well as its therapeutic potential in the treatment of cartilage-related diseases, such as osteoarthritis.

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