Cortical anchoring of the microtubule cytoskeleton is essential for neuron polarity.

He, Liu; Kooistra, Robbelien; Das, Ravi; Oudejans, Ellen; van Leen, Eric; Ziegler, Johannes; Portegies, Sybren; de Haan, Bart et al. · Elife · 2020

basic_science · Level V

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Abstract

The development of a polarized neuron relies on the selective transport of proteins to axons and dendrites. Although it is well known that the microtubule cytoskeleton has a central role in establishing neuronal polarity, how its specific organization is established and maintained is poorly understood. Using the in vivo model system <i>Caenorhabditis elegans</i>, we found that the highly conserved UNC-119 protein provides a link between the membrane-associated Ankyrin (UNC-44) and the microtubule-associated CRMP (UNC-33). Together they form a periodic membrane-associated complex that anchors axonal and dendritic microtubule bundles to the cortex. This anchoring is critical to maintain microtubule organization by opposing kinesin-1 powered microtubule sliding. Disturbing this molecular complex alters neuronal polarity and causes strong developmental defects of the nervous system leading to severely paralyzed animals.

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