Distinct mechanisms of the human mitoribosome recycling and antibiotic resistance.

Koripella, Ravi Kiran; Deep, Ayush; Agrawal, Ekansh K; Keshavan, Pooja; Banavali, Nilesh K; Agrawal, Rajendra K · Nat Commun · 2021

basic_science · Level V

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Abstract

Ribosomes are recycled for a new round of translation initiation by dissociation of ribosomal subunits, messenger RNA and transfer RNA from their translational post-termination complex. Here we present cryo-EM structures of the human 55S mitochondrial ribosome (mitoribosome) and the mitoribosomal large 39S subunit in complex with mitoribosome recycling factor (RRF<sub>mt</sub>) and a recycling-specific homolog of elongation factor G (EF-G2<sub>mt</sub>). These structures clarify an unusual role of a mitochondria-specific segment of RRF<sub>mt</sub>, identify the structural distinctions that confer functional specificity to EF-G2<sub>mt</sub>, and show that the deacylated tRNA remains with the dissociated 39S subunit, suggesting a distinct sequence of events in mitoribosome recycling. Furthermore, biochemical and structural analyses reveal that the molecular mechanism of antibiotic fusidic acid resistance for EF-G2<sub>mt</sub> is markedly different from that of mitochondrial elongation factor EF-G1<sub>mt</sub>, suggesting that the two human EF-G<sub>mt</sub>s have evolved diversely to negate the effect of a bacterial antibiotic.

Medical subject headings