Structural mechanism of tapasin-mediated MHC-I peptide loading in antigen presentation.

Jiang, Jiansheng; Taylor, Daniel K; Kim, Ellen J; Boyd, Lisa F; Ahmad, Javeed; Mage, Michael G; Truong, Hau V; Woodward, Claire H et al. · Nat Commun · 2022

basic_science · Level V

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Abstract

Loading of MHC-I molecules with peptide by the catalytic chaperone tapasin in the peptide loading complex plays a critical role in antigen presentation and immune recognition. Mechanistic insight has been hampered by the lack of detailed structural information concerning tapasin-MHC-I. We present here crystal structures of human tapasin complexed with the MHC-I molecule HLA-B*44:05, and with each of two anti-tapasin antibodies. The tapasin-stabilized peptide-receptive state of HLA-B*44:05 is characterized by distortion of the peptide binding groove and destabilization of the β<sub>2</sub>-microglobulin interaction, leading to release of peptide. Movements of the membrane proximal Ig-like domains of tapasin, HLA-B*44:05, and β<sub>2</sub>-microglobulin accompany the transition to a peptide-receptive state. Together this ensemble of crystal structures provides insights into a distinct mechanism of tapasin-mediated peptide exchange.

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