Critical soil moisture thresholds of plant water stress in terrestrial ecosystems.

Fu, Zheng; Ciais, Philippe; Feldman, Andrew F; Gentine, Pierre; Makowski, David; Prentice, I Colin; Stoy, Paul C; Bastos, Ana et al. · Sci Adv · 2022

basic_science · Level V

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Abstract

Plant water stress occurs at the point when soil moisture (SM) limits transpiration, defining a critical SM threshold (θ<sub>crit</sub>). Knowledge of the spatial distribution of θ<sub>crit</sub> is crucial for future projections of climate and water resources. Here, we use global eddy covariance observations to quantify θ<sub>crit</sub> and evaporative fraction (EF) regimes. Three canonical variables describe how EF is controlled by SM: the maximum EF (EF<sub>max</sub>), θ<sub>crit</sub>, and slope (S) between EF and SM. We find systematic differences of these three variables across biomes. Variation in θ<sub>crit</sub>, S, and EF<sub>max</sub> is mostly explained by soil texture, vapor pressure deficit, and precipitation, respectively, as well as vegetation structure. Dryland ecosystems tend to operate at low θ<sub>crit</sub> and show adaptation to water deficits. The negative relationship between θ<sub>crit</sub> and S indicates that dryland ecosystems minimize θ<sub>crit</sub> through mechanisms of sustained SM extraction and transport by xylem. Our results further suggest an optimal adaptation of local EF-SM response that maximizes growing-season evapotranspiration and photosynthesis.