iATPSnFR2: A high-dynamic-range fluorescent sensor for monitoring intracellular ATP.

Marvin, Jonathan S; Kokotos, Alexandros C; Kumar, Mukesh; Pulido, Camila; Tkachuk, Ariana N; Yao, Jocelyn Shuxin; Brown, Timothy A; Ryan, Timothy A · Proc Natl Acad Sci U S A · 2024

basic_science · Level V

Where this comes from

Abstract

We developed a significantly improved genetically encoded quantitative adenosine triphosphate (ATP) sensor to provide real-time dynamics of ATP levels in subcellular compartments. iATPSnFR2 is a variant of iATPSnFR1, a previously developed sensor that has circularly permuted superfolder green fluorescent protein (GFP) inserted between the ATP-binding helices of the <i>ε</i>-subunit of a bacterial F<sub>0</sub>-F<sub>1</sub> ATPase. Optimizing the linkers joining the two domains resulted in a ~fivefold to sixfold improvement in the dynamic range compared to the previous-generation sensor, with excellent discrimination against other analytes, and affinity variants varying from 4 µM to 500 µM. A chimeric version of this sensor fused to either the HaloTag protein or a suitable spectrally separated fluorescent protein provides an optional ratiometric readout allowing comparisons of ATP across cellular regions. Subcellular targeting the sensor to nerve terminals reveals previously uncharacterized single-synapse metabolic signatures, while targeting to the mitochondrial matrix allowed direct quantitative probing of oxidative phosphorylation dynamics.

Medical subject headings