m<sup>6</sup>A/IGF2BP3-driven serine biosynthesis fuels AML stemness and metabolic vulnerability.

Huang, Feng; Wang, Yushuai; Zhang, Xiuxin; Gao, Weiwei; Li, Jingwen; Yang, Ying; Mo, Hongjie; Prince, Emily et al. · Nat Commun · 2025

basic_science · Level V

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Abstract

Metabolic reprogramming of amino acids represents a vulnerability in cancer cells, yet the mechanisms underlying serine metabolism in acute myeloid leukemia (AML) and leukemia stem/initiating cells (LSCs/LICs) remain unclear. Here, we identify RNA N<sup>6</sup>-methyladenosine (m<sup>6</sup>A) modification as a key regulator of serine biosynthesis in AML. Using a CRISPR/Cas9 screen, we find that depletion of m<sup>6</sup>A regulators IGF2BP3 or METTL14 sensitizes AML cells to serine and glycine (SG) deprivation. IGF2BP3 recognizies m<sup>6</sup>A on mRNAs of key serine synthesis pathway (SSP) genes (e.g., ATF4, PHGDH, PSAT1), stabilizing these transcripts and sustaining serine production to meet the high metabolic demand of AML cells and LSCs/LICs. IGF2BP3 silencing combined with dietary SG restriction potently inhibits AML in vitro and in vivo, while its deletion spares normal hematopoiesis. Our findings reveal the critical role of m<sup>6</sup>A modification in the serine metabolic vulnerability of AML and highlight the IGF2BP3/m<sup>6</sup>A/SSP axis as a promising therapeutic target.

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