A male-pheromone-elevated transcription factor ZNF362.1 in female schistosomes determines sexual maturation.

Gu, Mengjie; Cheng, Wenjun; Li, Shan; Chen, Gongwen; Chen, Xu; Jiang, Ruiqi; Yuan, Minwei; Wang, Jing et al. · Sci Adv · 2026

basic_science · Level V

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Abstract

Egg production by female schistosomes drives both transmission and pathology of schistosomiasis, affecting over 200 million people. Female maturation relies on the male-derived pheromone β-alanyl-tryptamine (BATT), but underlying molecular mechanisms are unclear. We identified the BATT-responsive transcription factor gene <i>znf362</i> as a key regulator of female reproductive development. Functional studies showed that <i>znf362.1</i>, but not <i>znf362.2</i>, is essential for BATT-induced ovary and vitellaria maturation. Single-cell transcriptomics and in situ hybridization revealed up-regulation of <i>znf362.1</i> in oocytes and vitellaria S<sub>1</sub> cells after BATT exposure. Multiomics analysis showed ZNF362.1 directly activates <i>Smp_349410</i>, a female gonad-specific gene encoding a CPEB1 homolog. Loss of <i>znf362.1</i> or <i>Smp_349410</i> impaired oocyte and vitellocyte differentiation without affecting progenitors. Mechanistically, <i>Sm</i>CPEB1 promotes female ovary development by regulating polyadenylation of cyclin B1 mRNA and drives S<sub>1</sub> cell differentiation in the vitellaria. These findings define a transcriptional and post-transcriptional axis, BATT-<i>znf362.1</i>-<i>cpeb1</i>, that initiates female sexual maturation, offering mechanistic insight into schistosome reproduction and potential targets for schistosomiasis control.

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